Overview: The Biological Stakes of Megafire
Wildfire has shaped the biodiversity of North American landscapes for millions of years. Many of the continent's most distinctive and celebrated ecosystems — the giant sequoia groves of the Sierra Nevada, the chaparral shrublands of coastal California, the ponderosa pine savannas of the Pacific Northwest, the shortgrass prairies of the southern plains — evolved in intimate relationship with fire, and many species within them have life histories adapted to periodic burning. Seeds that germinate only after fire; bark structures evolved to resist low-intensity flames; post-fire successional habitats that support unique assemblages of cavity-nesting birds and pollinators: fire is not an external threat to these ecosystems but an intrinsic part of their ecological identity.
The wildfires documented in this database between 2015 and 2025, however, are fundamentally different in character from the fire regimes within which these species and ecosystems evolved. They burn larger, at higher severity, with longer intervals between consecutive burning events in some areas and at shorter intervals in others, and they arrive after a century of fire suppression has created fuel conditions with no historical parallel. The critical distinction — increasingly documented in the scientific literature — is between fire as an ecological process and fire as an ecological catastrophe: between the periodic, patchy, mostly low-to-moderate severity burns that sustained biodiversity for millennia, and the megafires that now kill virtually everything across hundreds of thousands of acres at a time.
A landmark 2023 study published in the Proceedings of the National Academy of Sciences by Ayars, Kramer, and Jones of the USDA Forest Service Rocky Mountain Research Station quantified the wildlife habitat impacts of California's unprecedented 2020–2021 megafire season. Their findings established the scale of the biological crisis: more than 19,000 km² of forest vegetation burned in just two years — ten times more than the historical average — potentially affecting the habitat of 508 vertebrate species. Of the more than 9,000 km² that burned at high severity, a full 89% occurred in large patches that exceeded historical estimates of maximum high-severity patch size. One hundred vertebrate species experienced fire across more than 10% of their geographic range. Sixteen of these were species of formal conservation concern under state or federal frameworks.
508 Vertebrate Species Affected
During the 2020–2021 California megafire season alone, the habitat of 508 vertebrate species was potentially impacted. One hundred of these species experienced fire across more than 10% of their entire geographic range — an unprecedented level of range-wide exposure for which "there is simply no frame of reference for understanding population responses," according to USDA Forest Service ecologist Gavin Jones. Sixteen species were formally listed as species of conservation concern. (Ayars, Kramer & Jones, PNAS, 2023)
The Architecture of Biodiversity Loss: How High-Severity Fire Destroys Life
To understand the biodiversity impacts of the fires reviewed in this database, it is essential to understand why high-severity fire — which kills the entire plant community, incinerates litter layers, sterilises surface soils, and eliminates standing dead wood — produces outcomes fundamentally different from those of lower-severity fire. At low-to-moderate severity, fire creates a mosaic of burned and unburned patches, opens the forest canopy to allow light-demanding plants to establish, removes accumulated fuel, and generates the biological diversity of structure — snags, downed logs, regenerating shrubs, mature trees — that supports a wide range of wildlife from cavity-nesting woodpeckers to large predators. This is the fire regime that shaped the species assemblages of western North American forests over thousands of years.
At the high severities now characteristic of megafires — where 75–100% of above-ground vegetation is killed across tens or hundreds of thousands of contiguous acres — the outcomes diverge dramatically. Large, homogeneous high-severity patches exceed the dispersal capacity of many species, creating vast dead zones that function as biological barriers rather than refugia. Seed banks in surface soils are destroyed by the heat of intense fires, eliminating the primary natural mechanism for post-fire plant community recovery. The soil mycorrhizal network — the fungal root-symbiosis system on which most forest tree species depend for survival — collapses. And without vegetation to anchor hillsides, post-fire erosion rapidly removes the thin layer of biologically productive topsoil that accumulated over centuries or millennia.
Fauna: Species by Species Impacts
The following profiles represent the best-documented cases of wildfire impact on individual animal species or species groups within the 2015–2025 review period. They are drawn from peer-reviewed research, U.S. Fish and Wildlife Service assessments, USDA Forest Service studies, and reports from conservation organisations with active field monitoring programmes.
🦉
Federally Threatened
Northern Spotted Owl
Strix occidentalis caurina
Dependent on old-growth and mature closed-canopy forest for nesting, roosting, and foraging. High-severity wildfire destroys this habitat structure. Research by Jones and Peery found that 22% of nesting sites used in 2014 were not reoccupied in 2015 after fires swept through nesting areas. Owls with GPS tracking tags were found to avoid burned areas larger than 100 hectares. The Bootleg Fire (2021, Oregon), McKinney Fire (2022), and Slater Complex (2020) all burned within critical spotted owl recovery habitat in the Klamath region. The species is already under severe pressure from barred owl competition and timber harvest.
Fires: Bootleg 2021 · McKinney 2022 · Slater 2020 · August Complex 2020
🦅
Federally Endangered
California Condor
Gymnogyps californianus
One of the world's rarest birds, with a global population of approximately 500–560 individuals (as of 2024), over half of which are in California. Condors require large territories with thermal updrafts and access to ungulate carcasses; they nest in cliff ledges and large cavities in old trees. The 2017 Thomas Fire, 2020 Creek Fire, 2021 Dixie Fire, and 2025 Dragon Bravo Fire all burned within or adjacent to established condor foraging and nesting range. Lead poisoning from carcasses of animals killed by fire (and shot by hunters with lead ammunition) creates a secondary mortality risk in post-fire landscapes.
Fires: Thomas 2017 · Creek 2020 · Dixie 2021 · Dragon Bravo 2025
🐟
Federally Threatened
Coho Salmon (Central CA & S. OR)
Oncorhynchus kisutch
Post-fire watershed contamination directly threatens Klamath River and coastal watershed coho salmon populations. Fires destroy streamside (riparian) vegetation that shades streams, maintaining cool temperatures required for salmon survival; post-fire erosion delivers fine sediment that smothers spawning gravels; increased peak flows associated with burned watersheds scour redds (nests) during critical incubation periods. The McKinney Fire (2022) and Slater-Devil Complex (2020) burned directly through critical Klamath River coho habitat. Central California coho populations were already reduced to fewer than 500 adults prior to the 2020 fire season.
Fires: McKinney 2022 · Slater 2020 · CZU Complex 2020
🐾
Proposed Endangered (CA)
California Spotted Owl
Strix occidentalis occidentalis
The USFWS proposed ESA listing for California Spotted Owl in 2023, explicitly citing wildfire as the "leading threat." Destructive megafires burned more spotted owl habitat in 2020 and 2021 alone than in the previous 35 years combined. The Sierra Nevada population requires forests with varying tree heights and a mix of open and dense areas; high-severity megafires replace this structural complexity with uniform burned-over landscapes. More than 1.7 million acres of Sierra Nevada forestland burned in wildfires since 2020. The KNP Complex, August Complex, Creek Fire, and Caldor Fire all burned within owl nesting territories.
Fires: KNP Complex 2021 · Creek 2020 · Dixie 2021 · Caldor 2021
🦡
State & Fed. Proposed Threatened
Pacific Fisher (S. Sierra Nevada)
Pekania pennanti
The Southern Sierra Nevada population of Pacific fisher is estimated at only 100–500 individuals — critically small. Fishers require large, structurally complex forests with abundant downed wood, dense canopy cover, and large-diameter trees for denning. Catastrophic wildfire — which removes precisely these structural elements — is identified by the USFWS as one of the biggest threats to the species. The 2021 KNP Complex Fire burned through known fisher habitat in Sequoia and Kings Canyon NPs. The Rough Fire (2015) burned 150,000 acres of Sequoia NF fisher habitat. The Dixie and Creek fires (2020–21) further fragmented the already tenuous population.
Fires: Rough 2015 · Creek 2020 · KNP Complex 2021 · Dixie 2021
🦦
Federally Endangered (Coastal DPS)
Humboldt Marten (Coastal Marten)
Martes caurina humboldtensis
One of North America's rarest carnivores, restricted to mature coastal forests of northern California and southern Oregon. Listed as Federally Endangered for its coastal distinct population segment in 2024. The species requires structurally complex old-growth and old-seral forests with dense understory — habitat that high-severity fire destroys. Save the Redwoods League notes the species "does well in the old-growth redwoods of the Northern California coast" — the precise habitat threatened by the CZU Lightning Complex (2020) and fires in the Klamath region.
Fires: CZU Complex 2020 · Slater Complex 2020 · McKinney 2022
🦤
Federally Threatened
Marbled Murrelet
Brachyramphus marmoratus
A seabird that nests exclusively on large moss-covered platforms in old-growth trees, often miles inland from the coast. Its unique nesting requirement means the species is entirely dependent on the continued existence of old-growth forest patches — precisely the habitat eliminated by high-severity fire. The CZU Lightning Complex (2020) burned 97% of Big Basin Redwoods State Park, which contains critical murrelet nesting habitat. The species faces additional pressure from corvid nest predation (jays and crows drawn to campgrounds in park areas) and is in steep, decades-long population decline.
Fires: CZU Complex 2020 · Thomas 2017 (coastal sycamore)
🐍
Species of Greatest Conservation Need
Long-Toed Salamander
Ambystoma macrodactylum
According to the USDA Forest Service / PNAS (2023) study by Ayars, Kramer, and Jones, the long-toed salamander experienced high-severity fire across a greater proportion of its geographic range than any other species examined — 14% of its total range burned at high severity in 2020–2021 alone. Research shows salamanders decline 1–2 decades post-fire in severely burned areas. As an amphibian with limited dispersal capacity, restricted habitat requirements (moist forest edges and wetlands), and low reproductive rate, the species faces serious long-term population impacts that may take decades to manifest.
Fires: 2020–2021 California Season (range-wide impact)
🐰
State Endangered (WA)
Washington Pygmy Rabbit
Brachylagus idahoensis
Following the 2020 Washington wildfire season — part of the Pacific Northwest fire siege that burned millions of acres — biologists estimated that fires had killed approximately 50% of the state's entire endangered pygmy rabbit population, leaving only about 50 individuals of North America's smallest rabbit species in Washington State. The animals inhabit sagebrush flats that burned during the 2020 fires. This near-total population-level mortality represents exactly the extinction-threat scenario conservation biologists warn of for range-restricted species. (Science/AAAS, September 2020)
Fires: 2020 Washington/Oregon wildfire season
🦃
Species of Conservation Concern
Greater Sage-Grouse
Centrocercus urophasianus
Following the 2020 Washington wildfire season, wildlife officials estimated that fires killed 30–70% of sage grouse and sharp-tailed grouse populations in affected areas. Sage-grouse are obligate sagebrush specialists that require mature sagebrush for cover, food, and nesting; fire destroys sagebrush and favours post-fire invasion by cheatgrass, which provides neither the cover nor the food quality that sage-grouse require. Sage-grouse populations across the western United States have been in long-term decline; the Smokehouse Creek Fire (2024, Texas) additionally affected lesser prairie chicken habitat in the Texas Panhandle.
Fires: 2020 WA season · Smokehouse Creek 2024
🦁
Species of Special Concern
Mountain Lion (Santa Monicas)
Puma concolor
The genetically isolated mountain lion population of the Santa Monica Mountains faces exceptional vulnerability. The 2018 Woolsey Fire burned 88% of Santa Monica Mountains NRA — nearly the entire territory of this population, estimated at 15–25 individuals. The famous P-22 (who ranged through Griffith Park) died in 2022, partly from the compounding stresses of fire-driven habitat loss and vehicle strikes. EcoSkills Academy estimates that the 2020 California fire season killed 300–600 mountain lions statewide — approximately 15% of the estimated total California mountain lion population. The 2025 Palisades Fire added further pressure on the Santa Monicas population.
Fires: Woolsey 2018 · 2020 CA season · Palisades 2025
🐿️
Federally Endangered (N. Rim endemic)
Kaibab Squirrel
Sciurus aberti kaibabensis
The Kaibab squirrel is entirely endemic to the Kaibab Plateau on the North Rim of Grand Canyon National Park — it exists nowhere else on Earth. Entirely dependent on Ponderosa pine forests for food (seeds, inner bark, fungi) and cover. The 2025 Dragon Bravo Fire burned extensively through the Kaibab Plateau's ponderosa pine forest, directly within this species' entire global range. The combination of an irreplaceable endemic population, a single-ecosystem dependency, and the scale of the 2025 fire creates acute extinction risk for this subspecies.
Fires: Dragon Bravo Fire 2025 (entire range burned)
Flora: Endangered Plants, Rare Species, and the Seed Bank Crisis
The impacts of high-severity wildfire on plant biodiversity are as severe as those on animal species, and in some respects more lasting, because plants are immobile: they cannot flee a fire, and if the individuals and seed banks of a rare species are eliminated from a location by fire, that species is gone from that site. California, which contains approximately 5,900 native plant species and ranks among the world's most botanically diverse regions, hosts hundreds of endemic species with restricted ranges — many of which exist in landscapes that have repeatedly burned during the 2015–2025 period.
Selected Fire-Impacted Rare & Endemic Flora
Giant Sequoia — Sequoiadendron giganteum2,000–3,200-year-old trees; 10% of world pop. killed 2020–21; not recoverable on human timescale
Coast Redwood — Sequoia sempervirensAncient trees in Big Basin SP survived CZU 2020 fire; visitor infrastructure destroyed; 97% of park burned
Coulter Pine — Pinus coulteriFire ecologist Camille Stevens-Rumann (CSU) notes this species with small ranges "could face trouble" from megafire; chaparral zone, Southern CA
Spalding's Catchfly — Silene spaldingiiFederally threatened perennial; fire-intolerant; threatened by invasive Ventenata grass (wiregrass) that creates extreme fire conditions in scablands of eastern OR
Santa Ana River Woolly Star — Eriastrum densifoliumFederally Endangered; small range in San Bernardino Valley; habitat adjacent to Woolsey (2018) and Palisades (2025) fire perimeters
Baker's Larkspur — Delphinium bakeriFederally Endangered; restricted to northern CA coast ranges; habitat intersects with fire perimeters of 2017 and 2020 North Bay region fires
Marsh Sandwort — Arenaria paludicolaFederally Endangered; one of California's rarest plants; coastal range habitat at risk from post-fire habitat conversion and invasive species post-fire colonisation
Native Sagebrush CommunitiesFire-intolerant; cheatgrass invasion post-fire prevents recovery; irreversible conversion of Great Basin sagebrush to annual grassland documented across OR, WA, NV following 2017–2021 fires
A particularly important — and frequently overlooked — dimension of fire impacts on plant biodiversity is the destruction of the soil seed bank: the reservoir of dormant seeds stored in the upper layers of soil that serves as the primary natural mechanism for post-fire vegetation recovery. High-severity fire, which generates soil temperatures exceeding 300–400°C at the surface, kills the vast majority of heat-sensitive seeds. For many native plants — including species with specific germination requirements that evolved in partnership with low-intensity fire — the destruction of the seed bank means that post-fire sites cannot recover to their pre-fire species composition without active human restoration, and in some cases cannot recover at all if the nearest seed sources are far outside the fire perimeter.
Research by Stevens-Rumann and colleagues has demonstrated that post-fire conifer regeneration failure is increasing substantially in the western United States, with a growing proportion of high-severity burn patches failing to recover as forest over 30+ year observation periods. In the Sierra Nevada, fire ecologists have documented the conversion of former conifer forest to shrubland in areas where high-severity fire exceeded the seed dispersal range of surviving trees. California's Office of Environmental Health Hazard Assessment notes that forest conversion to shrub or grassland has "adverse impacts on soil productivity, water quality, wildlife habitat, and carbon storage" that can persist for decades.
Soil Biodiversity: The Invisible Ecosystem
Beneath the visible destruction of above-ground plants and animals lies a biological catastrophe that receives far less attention: the collapse of soil biodiversity. Forest soils teem with bacteria, fungi, archaea, nematodes, mites, springtails, earthworms, and thousands of other organisms that together constitute the living foundation of terrestrial ecosystem function. A single tablespoon of healthy forest soil contains more microorganisms than there are humans on Earth. This underground community drives nutrient cycling, supports plant growth through mycorrhizal symbioses, regulates carbon storage, filters water, and maintains soil structure.
Research published in Nature Microbiology (2022) by scientists from Lawrence Berkeley National Laboratory characterised wildfire-driven changes in soil microbiomes across burn severity gradients in coniferous forests of Colorado and Wyoming. The study found severity-dependent losses of ectomycorrhizal fungi (EMF) — the root-symbiotic fungi on which almost all conifer tree species depend for nutrient acquisition and survival. Across the burn severity gradient, EMF relative abundance declined by a staggering 99%. The dominant EMF symbiont of lodgepole pine, Cenococcum geophilum, was completely absent from burned sites. Without EMF, planted conifer seedlings in post-fire reforestation projects face dramatically reduced survival rates, and natural tree regeneration is severely impaired.
A 2023 study in Science of the Total Environment examining prescribed fire versus wildfire impacts on soil microbiomes in California grasslands found that high-severity wildfire significantly reduced mycorrhizal fungal richness and colonisation rates — with direct downstream consequences for native plant recovery and the displacement of mycorrhizal-dependent native species by non-mycorrhizal invasive plants that can establish more easily on mycorrhizal-depleted soils.
Invasive Species and the Post-Fire Biodiversity Crisis
High-severity wildfire creates ideal conditions for invasive species establishment. Burned landscapes offer open canopies, disturbed soils, reduced competition from perennial native plants, and altered soil chemistry that many invasive species are far better equipped to exploit than the native species they displace. The cheatgrass (Bromus tectorum) invasion of the Great Basin is the most studied example: this annual grass from Eurasia thrives in post-fire conditions, establishes dense monocultures that suppress native sagebrush and bunchgrass regeneration, and creates fine fuel beds that burn at short intervals — creating a self-reinforcing cycle of fire, invasive dominance, and loss of native biodiversity that has permanently transformed millions of acres of the interior West.
In Hawaii, the 2023 Lahaina fire was directly enabled by the invasion of non-native grasses including buffelgrass (Cenchrus ciliaris) and molasses grass (Melinis minutiflora), which replaced native dryland forest and created continuous, highly flammable fuel beds across the landscape. These grasses are now the dominant vegetation across large areas of Maui's lowlands, and their continued expansion virtually guarantees recurring and worsening fire conditions that threaten the remaining fragments of native Hawaiian dryland forest — one of the most imperilled plant communities on Earth, home to dozens of endemic Hawaiian plant species not found anywhere else in the world.
In the Pacific Northwest, the invasive wiregrass (Ventenata dubia) is emerging as a new threat, colonising the cool, high-elevation scablands of eastern Oregon and Washington that were previously beyond the range of cheatgrass. A post-fire study of the Corner Creek Fire found wiregrass creating fuel loads 50 times greater than wiregrass-free areas — and threatening the rare endemic Spalding's catchfly (Silene spaldingii), a federally protected perennial of the rocky scabland habitat.
Future Wildfire Risk for Global Biodiversity
A major 2025 study published in Nature Climate Change modelled future wildfire exposure for all non-marine species identified as threatened by increased fire frequency or intensity globally. It found that under SSP2-4.5 (a moderate emissions scenario), 83.9% of the 9,592 fire-vulnerable species identified would face higher wildfire risk by the end of the century. In the western United States specifically, fire season duration is projected to more than double in high-latitude and high-elevation regions. The study's conclusion is stark: wildfire is becoming a primary driver of global biodiversity loss, operating in synergy with habitat destruction, invasive species, and climate change to create compound threats that individually threatened species may be unable to survive.
| Species |
Status |
Primary Fire Threat |
Key Fires Impacted (2015–2025) |
Conservation Prognosis |
| Northern Spotted Owl |
Fed. Threatened |
Old-growth habitat loss |
Bootleg 2021, McKinney 2022, August Complex 2020 |
Critical; barred owl competition adds pressure |
| California Condor |
Fed. Endangered |
Foraging range destruction; lead in carcasses |
Thomas 2017, Creek 2020, Dixie 2021, Dragon Bravo 2025 |
Guarded; active recovery programme ongoing |
| Coho Salmon (Central CA/S. OR) |
Fed. Threatened |
Watershed contamination; stream heating; sediment |
McKinney 2022, Slater 2020, CZU 2020 |
Critical; populations already near extirpation |
| California Spotted Owl |
Proposed ESA Listing 2023 |
Sierra Nevada habitat megafire destruction |
Creek 2020, KNP Complex 2021, Dixie 2021, Caldor 2021 |
Critical; 35 yrs of habitat lost in 2 years |
| Pacific Fisher (S. Sierra Nevada) |
Fed. Proposed Threatened |
Forest structure loss; denning habitat |
Rough 2015, Creek 2020, KNP Complex 2021, Dixie 2021 |
Critical; 100–500 individuals remaining |
| Humboldt Marten |
Fed. Endangered (coastal DPS) |
Old-growth coastal forest destruction |
CZU Complex 2020, Slater 2020, McKinney 2022 |
Critical; extreme range restriction |
| Marbled Murrelet |
Fed. Threatened |
Old-growth nest platform tree loss |
CZU Complex 2020, Thomas 2017 |
Seriously declining; decades-long downward trend |
| Long-Toed Salamander |
Species of Greatest Conservation Need |
14% of total range burned at high severity (2020–21) |
2020–21 CA megafire season |
Unknown; no historical baseline for this scale |
| Washington Pygmy Rabbit |
State Endangered (WA) |
Sagebrush habitat incinerated |
2020 WA/OR fire season |
Critical; ~50% of population killed in 2020 |
| Giant Sequoia |
State Endangered (CA, 2021 emergency) |
Megafire kills ancient trees; 10% world pop. lost |
Castle Fire 2020, KNP Complex 2021, Windy Fire 2021 |
Irreversible; trees 2,000–3,200 yrs old; no replacement |
| Kaibab Squirrel |
Fed. Endangered (N. Rim endemic) |
Entire range on Kaibab Plateau burned (2025) |
Dragon Bravo Fire 2025 |
Acute extinction risk; entire global range affected |
| Lesser Prairie Chicken (TX) |
Fed. Threatened |
Texas Panhandle grassland habitat incinerated |
Smokehouse Creek 2024 |
Serious concern; habitat reduced further |
The Biodiversity Value of What Has Already Been Lost
Attempts to place economic value on biodiversity loss from wildfire face profound methodological and philosophical challenges. Unlike a burned house, which can be rebuilt, the genetic and evolutionary heritage of an extinct species — or the irreplaceable mycorrhizal community of an ancient forest soil — has no replacement cost. The giant sequoias killed in the 2020–2021 California fires had grown for between 2,000 and 3,200 years. No amount of conservation investment can accelerate their replacement; the youngest sequoia planted today will not reach reproductive maturity for decades and will not approach the ecological significance of a 2,000-year-old tree for two thousand years.
Research published in Nature Climate Change (2025) has begun to develop frameworks for quantifying the future biodiversity cost of fire-driven habitat loss, using IUCN Red List assessments and species distribution models to project extinction risk trajectories under different fire scenario. The key finding is that biodiversity losses from fire are not merely acute — they have long tails. A species whose population is reduced by 50% in a single fire season does not necessarily recover in the next decade; many small-population species become caught in the extinction vortex of demographic and genetic stochasticity that makes recovery increasingly improbable as population size falls.
The convergence of wildfire intensification with other biodiversity stressors — habitat fragmentation, climate warming, drought, invasive species, and pollution — creates compound threats whose combined severity likely exceeds the sum of their individual parts. The northern spotted owl faces not only wildfire but also competition from the aggressively expanding barred owl; coho salmon face not only wildfire-driven watershed contamination but also warming stream temperatures, water diversion, and oceanic ecosystem changes; Pacific fishers face not only habitat loss from fire but also the legacy of historic trapping, rodenticide exposure from cannabis cultivation sites, and road mortality. For already-imperilled species, each new fire season brings compound threats to populations that may already be below the minimum viability threshold.
Bibliography & References
[1]Gajendiran, K., Kandasamy, S., and Narayanan, M. (2024). "Influences of wildfire on the forest ecosystem and climate change: A comprehensive study." Environmental Research, 243, 117851. DOI: 10.1016/j.envres.2023.117851. https://pubmed.ncbi.nlm.nih.gov. See also: Defenders of Wildlife. (April 2025). "Animals in the Aftermath: Southern Sierra Nevada Fishers After Wildfires." https://defenders.org
[2]California Wildfire & Forest Resilience Task Force. (2023). "Central California Regional Profile: Biodiversity Conservation." https://wildfiretaskforce.org. See also: California Office of Environmental Health Hazard Assessment (OEHHA). (2022). Indicators of Climate Change in California: Wildfires. https://oehha.ca.gov
[3]Ayars, J., Kramer, H.A., and Jones, G.M. (2023). "The 2020 to 2021 California megafires and their impacts on wildlife habitat." Proceedings of the National Academy of Sciences, 120(48), e2312909120. DOI: 10.1073/pnas.2312909120. https://pmc.ncbi.nlm.nih.gov. See also: U.S. Forest Service. (March 2024). "Extreme Wildfires Take a Toll on California Wildlife." https://www.fs.usda.gov
[4]Davis, K.T. et al. (2019). "Wildfires and climate change push low-elevation forests across a critical environmental threshold, triggering a major loss of forest cover." PNAS, 116(13). DOI: 10.1073/pnas.1815107116. See also: Journal of Student Research. (2022). "California in Flames: A Literature Review on the Causes and Effects of Wildfires." https://www.jsr.org
[5]California Department of Fish and Wildlife. (2026). "Every animal that's endangered or threatened in California in 2026." https://californiatoday.com. See also: Almanac.com. (January 2026). "Endangered Species of Plants." https://www.almanac.com
[6]Stevens-Rumann, C. et al. (AAAS/Science, 2020). Fire ecologist commentary cited in: Voosen, P. (September 30, 2020). "As wildfires continue in western United States, biologists fear for vulnerable species." Science/AAAS. https://www.science.org
[7]California OEHHA. (2022). Indicators of Climate Change in California: Wildfires Chapter. Citing: Shive, K.L. et al. (2022). "Ancient trees and modern wildfires: Declining resilience to wildfire in the highly fire-adapted giant sequoia." Forest Ecology and Management, 511, 120110. See also: Smith, J.L. (2000). Wildland fire in ecosystems: effects of fire on fauna. USDA Forest Service RMRS-GTR-42-vol. 1.
[8]Whitman, T. et al. (2022). "Wildfire-dependent changes in soil microbiome diversity and function." Nature Microbiology, 7, 1419–1430. DOI: 10.1038/s41564-022-01203-y. https://pmc.ncbi.nlm.nih.gov. See also: USDA Forest Service Research and Development. (2022). Wildfire-dependent changes in soil microbiome. https://research.fs.usda.gov
[9]Emam, T. et al. (2023). "Prescribed versus wildfire impacts on exotic plants and soil microbes in California grasslands." Applied Soil Ecology, 182. DOI: 10.1016/j.apsoil.2022.104715. https://www.sciencedirect.com. See also: Caiafa et al. (2025). "Wildfire impact on soil microbiome life history traits and roles in ecosystem carbon cycling." ISME Communications. https://pmc.ncbi.nlm.nih.gov
[10]Balch, J.K. et al. (2013). "Human-started wildfires expand the fire niche across the United States." PNAS. See also: arXiv. (2020). "Invasive species, extreme fire risk, and toxin release under a changing climate." DOI: arXiv:2008.01035. https://arxiv.org
[11]Wikipedia. (2025). "2023 Hawaii Wildfires." [Invasive grass fuel load section.] https://en.wikipedia.org. See also: ClimatTalk. (October 2024). "Wildfires: Their Impact on Climate, Biodiversity and Society." https://climatalk.org
[12]Pennisi, E. (Science/AAAS, 2022). "Flammable invasive grasses are increasing risk of devastating wildfires." Science. [Wiregrass / Spalding's catchfly section.] https://www.science.org
[13]Ruffault, J. et al. (2025). "Wildfire risk for species under climate change." Nature Climate Change. DOI: 10.1038/s41558-026-02600-5. https://www.nature.com
[14]National Park Service. (November 2021). "Giant Sequoia Mortality Estimates Released for the 2021 KNP Complex and Windy Fire." https://www.nps.gov. See also: Sierra Club. (2022). "A Year in the Life of a Burning Forest." https://www.sierraclub.org
[15]U.S. Fish & Wildlife Service. (June 2022). "Examining the Impacts of Wildfire on Fishers." https://www.fws.gov. See also: USFWS. (2023). "3 Efforts Underway that Support the California Spotted Owl and Reduce Wildfire Risk." https://www.fws.gov. See also: Audubon Society. (2025). "Fire Is a Major Threat to California Spotted Owls—Could it Also Help Save Them?" https://www.audubon.org
[16]EcoSkills Academy. (January 2025). "California Wildfires: Wildlife Habitat Under Attack." https://ecoskills.academy. See also: Curlewcall.org. (November 2025). "5 Major Threats to Biodiversity in America." https://www.curlewcall.org
[17]Save the Redwoods League. (July 2022). "Threatened and endangered wildlife in the redwoods." https://www.savetheredwoods.org
[18]Voosen, P. (September 30, 2020). "As wildfires continue in western United States, biologists fear for vulnerable species." Science/AAAS. [Pygmy rabbit, sage grouse, spotted owl sections.] https://www.science.org
[19]Courthouse News Service. (November 2023). "Researchers still hazy on effect of intense wildfires on California wildlife." Quoting USDA Forest Service ecologist Gavin Jones. https://www.courthousenews.com
[20]Ward, G. et al. (2021). "Resilience of terrestrial and aquatic fauna to historical and future wildfire regimes in western North America." Ecology and Evolution. DOI: 10.1002/ece3.8091. https://pmc.ncbi.nlm.nih.gov
[21]Hoffman, K.M. et al. (2021). "Conservation of Earth's biodiversity is embedded in indigenous fire stewardship." Proceedings of the National Academy of Sciences, 118(32). DOI: 10.1073/pnas.2105073118. Cited in: Journal of Student Research. (2022). California in Flames. https://www.jsr.org